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1. ANAXAGOREA A. St.-Hil., Nouv. Bull. Sci. Soc. Philom. Paris 1825: 91. 1825.

by Julian A. Steyermark and Paul E. Berry

Shrubs or small to medium-sized trees. Leaves chartaceous or subcoriaceous. Inflorescence axillary, sometimes leaf-opposed or terminal, occasionally cauliflorous, shortly pedunculate or epedunculate; lower bracts often caducous; upper bracts usually borne near flower, caducous or not. Flower buds globose, ovoid, or conical; flowers bisexual, fleshy, usually not opening widely, with brown, simple or stellate, microscopic hairs. Sepals valvate to imbricate, ovate to broadly ovate, free or connate, shorter than the petals; petals valvate, usually fleshy, green, white, yellowish, or red, the outer 3 somewhat or conspicuously longer than the inner 3. Stamens 10--200, apical prolongation of connective flat, triangular or quadrangular, innermost stamens staminodial; anthers not locellate. Carpels 5--45, each with 2 subbasal ovules. Monocarps free, clavate to dolabriform (hatchet- or golf club-shaped), basally stipitate, apically beaked, dehiscent along the ventral suture. Seeds 2 per monocarp, shiny, black, without an aril.

Mexico, Central America, Colombia, Venezuela, Trinidad, Guyana, Suriname, French Guiana, Ecuador, Peru, Brazil, Bolivia, tropical Asia; ca. 26 species, 9 species in Venezuela, all in the flora area.

Key to the Species of Anaxagorea

1. Leaves 25--75 × 7--25 cm, secondary veins 18--28 on each side of the mid-vein; petioles 1.5--4.5(--6) cm long, 4--8(--10) mm diameter...A. gigantophylla

1. Leaves and petioles usually smaller than above and the secondary veins fewer...2

2(1). Leaves narrowly to very narrowly elliptic to elliptic-ovate, 5--16 × 0.5--3.5 cm...3

2. Leaves broader and/or longer than above...4

3(2). Leaves 6--16 × 1.5--3.5 cm, midvein on upper surface slightly impressed; outer petals with a thick keel on the inner surface...A. angustifolia

3. Leaves very narrow, 5--8 × 0.5--1 cm, midvein on upper surface flat, not impressed; outer petals not keeled on inner surface...A. rheophytica

4(2). Midvein of the upper surface of leaf distinctly raised or convex; young twigs, perianth, and fruits sparsely puberulous to glabrous; fruits yellow to green with reddish tinge (when live)...A. brachycarpa

4. Midvein of the upper surface of leaf impressed or flat, or if somewhat elevated then young twigs, perianth, and fruits densely puberulous; fruits usually darker colored or with brown or reddish pubescence...5

5(4). Pedicels (of individual flowers, not the inflorescence rachis) 15--32 mm long, the upper bracts persistent until shortly after anthesis and situated (3--)5--15 mm below the calyx, about 1/3--1/2 way down the pedicel; petioles 7--21 mm long...A. petiolata

5. Pedicels usually < 15 mm long, the upper bracts more readily caducous and borne closer to the calyx than above; petioles usually shorter than above...6

6(5). Midvein of the upper surface of leaves ³ 1.5 mm broad at the base, often somewhat raised; hairs on petioles and perianth dark purplish red...A. rufa

6. Midvein of the upper surface of leaves < 1.5 mm broad at the base, flat to usually impressed; hairs on petioles and perianth brown or brownish red at most...7

7(6). Inflorescence axillary, cauliflorous, or borne at the shoot apex; sepals erect; outer petals keeled on the inner surface; vegetative parts usu-ally glabrous...A. brevipes

7. Inflorescence axillary or cauliflorous, but not borne at the shoot apex; sepals erect to recurved; outer petals not keeled on the inner surface; vegetative parts generally with fine brown or brownish red pubescence...8

8(7). Sepals usually £ 5 mm long, caducous or if persistent then spreading to upwardly curving; outer petals 7--12 mm long... A. acuminata

8. Sepals 6--13 mm long, often persistent and becoming recurved to re-flexed; outer petals 10--18(--23) mm long...A. dolichocarpa