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Michael Grayum 9660 Det. by: A. R. Smith, 1992 Phenology: 16 February 1990 09:54N 84:17W 900 m SAN JOSE, MORA Z.P. El Rodeo Valle del Tárcoles. Terrestrial ferns, occasional in primary forest understory. Rhizome erect, briefly prolonged (to 10-15 cm above ground). Fertile fronds narrower and on relatively longer petioles than sterile fronds.