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Michael Grayum 10207 Det. by: A. R. Smith, 1994 Phenology: 29 February 1992 10:18N 84:44W 850-900 m ALAJUELA, SAN RAMON R.B. Monteverde Cordillera de Tilarán. Terrestrial on damp, shaded slope near trail. Rhizome stout, erect. Note buds in axils of proximal pinnae on both sterile and fertile fronds.
Michael Grayum 9049 Det. by: A. R. Smith, 1993 Phenology: 17 November 1988 10:49N 84:53W 80 m ALAJUELA, UPALA Llanura de Guatuso. Tall, coarse fern, locally abundant on sandy shore of river. Rhizome very stout, creeping.